They lived more than two million years ago and almost 700 000 years apart. They belonged to the same species and they have finally been reunited at Maropeng a'Africa at the Cradle of Humankind.
In what academics have described as an internationally historic event, the skull of Mrs â€¦ Mr â€¦ or Ms Ples (the gender has not been agreed on) and the Taung child will be on view for the first time to the public at the new exhibition centre.
Ples is believed to be about two million years old and the four-year-old child about 2,7-million years old. Ples's skull has been around the world and examined by many palaeontologists.
Dr Francis Thackeray, curator of the Transvaal Museum, said on Tuesday it was "a rare occasion that the public will get to see the original fossil".
The skull will be on display until May 14, together with 50 fossils called the Original Fossil Display.